Teachers are the most important school-specific factor in student learning. Yet, little evidence exists linking teacher professional development programs and the strategies or activities that comprise them to student achievement. In this paper, we examine a fellowship model for professional development designed and implemented by Leading Educators, a national nonprofit organization that aims to bridge research and practice to improve instructional quality and accelerate learning across school systems. During the 2015-16 and 2016-17 school years, Leading Educators conducted its fellowship program for two cohorts of teachers and school leaders to provide these educators ongoing, collaborative, job-embedded professional development and to improve student achievement. Relying on quasi-experimental methods, we find that a school’s participation in the fellowship program significantly increased student proficiency rates in English language arts and math on state achievement exams. Student achievement benefitted from a more sustained duration of participation in the fellowship program, varied depending on the share of a school’s educators who participated in the fellowship, and differed based on whether fellows independently selected into the program or were appointed to participate by their school leaders. Taken together, findings from this paper should inform professional learning organizations, schools, and policymakers on the design, implementation, and impact of educator professional development.